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When Dr Harper Avery, the namesake of the Avery award, is rushed to the hospital, his unusual request for surgery heightens tension between Derek and Richard. The doctors are surprised to learn that Jackson is the medical legend’s grandson. Meanwhile, Callie and Arizona help Mark and Teddy move on from their rocky past relationships.
The team tries to diagnose a famous blogger but discover that her insistence on communicating on the Internet about her case interferes with their ability to cure her. Meanwhile, House and Wilson each discover secrets about each other, and Chase is convinced to go on a date.
Izzie spends time with a fellow cancer patient and contemplates whether or not to undergo a risky surgery, as Mark is ready to take his relationship with Lexie to the next level and a patient makes Owen reassess his place at Seattle Grace. George delivers stunning news to Bailey, sending shockwaves throughout the hospital, and Izzie’s friends anxiously await her recovery from surgery. Meanwhile Bailey is surprisingly displeased after being accepted into the pediatric fellowship program, and the victim of a near-fatal traffic accident brings the talents of Seattle Grace’s doctors together.
As Meredith, Derek and all of their friends prepare for the big wedding, a group of college students, injured in a car crash as they were heading to their graduation ceremony, is admitted to the ER. Meanwhile, Izzie fears the worst when she begins experiencing familiar symptoms.
As preparations continue for Meredith and Derek’s wedding, several residents must face the arrival of their parents at the hospital: Izzie’s mother Robbie causes a stir when she pays a surprise visit, and Thatcher returns to make amends with Meredith and Lexie. The Chief clashes with Meredith over a controversial medical case.